Qassem Aqlan was shot dead by gunmen on a motorbike as he drove along Sittin Street, a main artery of the Yemeni capital, the official said, adding the assailants managed to escape.
A US diplomat said Aqlan was in charge of coordinating a US and Yemeni probe into an attack last month on the embassy by protesters angry over an anti-Islam film that triggered a wave of deadly demonstrations across the Muslim world.
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